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Support for third-party writing assistance for this article, provided by Aditi Mehta, MSc, Barney Johnson, BA, and Sona Popat, BA, from Costello Medical, London, UK, was funded by UCB in accordance with Good Publication Practice (GPP 2022) guidelines

doi: 10.1073/pnas.0600445103 96 TanLWangJTanizakiJHuangZArefARRusanMet al

Supplementation with GSH is generally ineffective at increasing cellular glutathione levels because most human cells cannot uptake extracellular GSH due to the lack of the membrane-bound ectoenzyme -glutamyltranspeptidase (-GT) or due to an unfavorable concentration gradient across the cell membrane 10,16

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